西北植物学报
西北植物學報
서북식물학보
ACTA BOTANICA BOREALI-OCCIDENTALIA SINICA
2001年
3期
546-550
,共5页
曹玉芳%辛华%初庆刚%吕瑞云%陶世蓉
曹玉芳%辛華%初慶剛%呂瑞雲%陶世蓉
조옥방%신화%초경강%려서운%도세용
侧柏%小孢子囊%绒毡层%发育
側柏%小孢子囊%絨氈層%髮育
측백%소포자낭%융전층%발육
侧柏[Platycladus orientalis (L.) Franco]小孢子囊壁包括3层细胞:表皮、中层和绒毡层。中层细胞为1层扁平的细胞。绒毡层细胞属于分泌型。成熟的绒毡层细胞除了有单核和双核细胞外,还有三核和四核等多核细胞。细胞核有圆形和长椭圆形2种形态。绒毡层细胞的染色质伴随着小孢子母细胞减数分裂有一个浓缩和伸展的时期,这个时期影响营养物质向小孢子囊内部转运。绒毡层细胞发育的初期就为造孢细胞提供营养,后期解体时,分泌的乌氏体不是散乱地而是有组织地向花粉粒的表面转移。中层和绒毡层细胞最终作为营养被全部吸收利用。
側柏[Platycladus orientalis (L.) Franco]小孢子囊壁包括3層細胞:錶皮、中層和絨氈層。中層細胞為1層扁平的細胞。絨氈層細胞屬于分泌型。成熟的絨氈層細胞除瞭有單覈和雙覈細胞外,還有三覈和四覈等多覈細胞。細胞覈有圓形和長橢圓形2種形態。絨氈層細胞的染色質伴隨著小孢子母細胞減數分裂有一箇濃縮和伸展的時期,這箇時期影響營養物質嚮小孢子囊內部轉運。絨氈層細胞髮育的初期就為造孢細胞提供營養,後期解體時,分泌的烏氏體不是散亂地而是有組織地嚮花粉粒的錶麵轉移。中層和絨氈層細胞最終作為營養被全部吸收利用。
측백[Platycladus orientalis (L.) Franco]소포자낭벽포괄3층세포:표피、중층화융전층。중층세포위1층편평적세포。융전층세포속우분비형。성숙적융전층세포제료유단핵화쌍핵세포외,환유삼핵화사핵등다핵세포。세포핵유원형화장타원형2충형태。융전층세포적염색질반수착소포자모세포감수분렬유일개농축화신전적시기,저개시기영향영양물질향소포자낭내부전운。융전층세포발육적초기취위조포세포제공영양,후기해체시,분비적오씨체불시산란지이시유조직지향화분립적표면전이。중층화융전층세포최종작위영양피전부흡수이용。
The microsporangium wall cell of Platycladus orientalis (L.) Franco consists of three layers:the epidermis,the middle layer and the tapetum.The form of middle layer cell is flat.The tapetum is secretery type.There are tri-and tetra-caryotic cells in mature tapetum as well as mono-and di-caryotic cells.The form of tapetal cell nucleus is round or long elliptical.The reason of the formation of polycaryotic cell is that the nuclear division of dicaryotic tapetal cell.The chromatin of tapetal cell has a period of becoming condensed and then spreading accompanying the meiosis of the microspore mother cells,which will affect the entering of nutrient into the inner part of microsporangium.The tapetal cell has under taken the task of providing nutrition for the sporogenous cells at earlier development.Ubisch body secreted from tapetal cell later transport to the surface of pollen grain not in scattered but in organized form.The middle layer and the tapetum cells are totally absorbed as nutrient at last.