植物学报
植物學報
식물학보
ACTA BOTANICA SINICA
2003年
1期
15-22
,共8页
华夏植物群%早二叠世%煤核%美丽"鳞木"%茎%解剖构造
華夏植物群%早二疊世%煤覈%美麗"鱗木"%莖%解剖構造
화하식물군%조이첩세%매핵%미려"린목"%경%해부구조
Cathaysian Flora%Early permian%coal balls%"Lepidodendron" pulchrum%stems%anatomy
描述了山东兖州矿区太原组(早二叠世早期)煤核中一种具叶座的鳞木类茎的解剖构造.叶座呈略不对称的菱形,下侧边略长于上侧边,顶、底角均平截,底角并略呈拖延状;叶座上部突起较强烈,叶痕双凸镜形,叶迹呈宽V字形,叶迹与侧通气束可能都以近水平状在叶座内向外延伸;叶舌穴深,自叶座最内处向外斜伸,开口位于叶痕顶角附近.叶座底部表面具横皱纹.茎可能具管状中柱和薄壁组织的髓.初生木质部外始式,外缘较平滑.仅见有外部皮层,由径向伸长的壁较厚的细胞条带和夹于其间的空腔交替排列构成,空腔内可见弧形或宽V字形叶迹,其凹面朝向内方.周皮较发育,由近等量发育的木栓层和栓内层组成.经比较,当前标本与华夏植物区的印痕--压型化石种美丽鳞木在叶座的形态上非常相似,暂归入该种.由于无论是当前标本还是美丽鳞木的模式标本与狭义的鳞木属都有很大区别,因此美丽"鳞木"的确切归属还有待于今后进一步对保存更好的具解剖构造的茎和生殖器官的研究.美丽"鳞木"与欧美植物区几个乔木状鳞木类植物属的茎都不完全相同,具有它们的混合特征,很可能代表了一个新属.美丽"鳞木"是目前华夏植物区研究得最详细的一种具解剖构造的鳞木类的茎,对于研究华夏植物区鳞木类植物的起源和演化具有一定的意义.
描述瞭山東兗州礦區太原組(早二疊世早期)煤覈中一種具葉座的鱗木類莖的解剖構造.葉座呈略不對稱的蔆形,下側邊略長于上側邊,頂、底角均平截,底角併略呈拖延狀;葉座上部突起較彊烈,葉痕雙凸鏡形,葉跡呈寬V字形,葉跡與側通氣束可能都以近水平狀在葉座內嚮外延伸;葉舌穴深,自葉座最內處嚮外斜伸,開口位于葉痕頂角附近.葉座底部錶麵具橫皺紋.莖可能具管狀中柱和薄壁組織的髓.初生木質部外始式,外緣較平滑.僅見有外部皮層,由徑嚮伸長的壁較厚的細胞條帶和夾于其間的空腔交替排列構成,空腔內可見弧形或寬V字形葉跡,其凹麵朝嚮內方.週皮較髮育,由近等量髮育的木栓層和栓內層組成.經比較,噹前標本與華夏植物區的印痕--壓型化石種美麗鱗木在葉座的形態上非常相似,暫歸入該種.由于無論是噹前標本還是美麗鱗木的模式標本與狹義的鱗木屬都有很大區彆,因此美麗"鱗木"的確切歸屬還有待于今後進一步對保存更好的具解剖構造的莖和生殖器官的研究.美麗"鱗木"與歐美植物區幾箇喬木狀鱗木類植物屬的莖都不完全相同,具有它們的混閤特徵,很可能代錶瞭一箇新屬.美麗"鱗木"是目前華夏植物區研究得最詳細的一種具解剖構造的鱗木類的莖,對于研究華夏植物區鱗木類植物的起源和縯化具有一定的意義.
묘술료산동연주광구태원조(조이첩세조기)매핵중일충구협좌적린목류경적해부구조.협좌정략불대칭적릉형,하측변략장우상측변,정、저각균평절,저각병략정타연상;협좌상부돌기교강렬,협흔쌍철경형,협적정관V자형,협적여측통기속가능도이근수평상재협좌내향외연신;협설혈심,자협좌최내처향외사신,개구위우협흔정각부근.협좌저부표면구횡추문.경가능구관상중주화박벽조직적수.초생목질부외시식,외연교평활.부견유외부피층,유경향신장적벽교후적세포조대화협우기간적공강교체배렬구성,공강내가견호형혹관V자형협적,기요면조향내방.주피교발육,유근등량발육적목전층화전내층조성.경비교,당전표본여화하식물구적인흔--압형화석충미려린목재협좌적형태상비상상사,잠귀입해충.유우무론시당전표본환시미려린목적모식표본여협의적린목속도유흔대구별,인차미려"린목"적학절귀속환유대우금후진일보대보존경호적구해부구조적경화생식기관적연구.미려"린목"여구미식물구궤개교목상린목류식물속적경도불완전상동,구유타문적혼합특정,흔가능대표료일개신속.미려"린목"시목전화하식물구연구득최상세적일충구해부구조적린목류적경,대우연구화하식물구린목류식물적기원화연화구유일정적의의.
This study reports the anatomical structures of a kind of lepidodendralean stem in coal balls from the early Early Permian Taiyuan Formation in Yanzhou Mining District, southwestern Shandong Province,North China. The leaf cushion is slightly rhomboid in outline with a height of 9 - 10 mm and a width of 7.5 -9.0 mm and its lower sides are slightly longer than the upper ones. The top and basic angles of the leaf cushion are truncate and the basic angle is slightly elongate. The upper part of the leaf cushion is strongly highrising. The leaf scar is large and lenticular in shape. The leaf trace is wide and V-shaped in the leaf scar, and horizontally elongate within the leaf cushion. The leaf trace and lateral parichnos strand extend at a nearly horizontal course outward within the leaf cushion. The ligular pit is deep and extends outward at an oblique course and its aperture is located near the top angle of the leaf scar. No infrafoliar parichnos strands are present. The stem is probably siphonostelic and its pith is probably parenehymatous. The primary xylem is exarch with a nearly smooth outer margin. Only the outer cortex is present and it consists of alternately-arranged radial cell bands and gaps within which the are-shaped or V-shaped leaf traces can be seen. The concave side of the leaf trace is toward the center of the stem. No bundle sheath is developed. Periderm is well-developed and consists of phelloderm and phellem in nearly equal thickness. Compared with the lepidodendralean stems of the Cathaysian and Euramerican Floras, the present specimens are most close to an impression-compression species Lepidodendron pulchrum Zhang in morphology of the leaf cushion and they are put into this species temporarily. Whether the present specimens or the type specimens of L. pulchrum are very different from Lepidodendron Stemburg sensu DiMichele, thus the correct nomenclature and classification of L. pulchrum needs to be reconsidered based on the study of better- and anatomically-preserved stems and fertile organs in the future.Because "L". pulchrum possesses the mixed features of several genera of Euramerican lepidodendralean stems, it bears significance to study the origin and evolution of the Cathaysian lepidodendralean lycopods.