江西农业学报
江西農業學報
강서농업학보
ACTA AGRICULTURAE JIANGXI
2015年
8期
25-29,35
,共6页
付娟%李霞%丁君辉%唐启源%王若仲
付娟%李霞%丁君輝%唐啟源%王若仲
부연%리하%정군휘%당계원%왕약중
水稻%生长锥%幼穗%维管束%形态
水稻%生長錐%幼穗%維管束%形態
수도%생장추%유수%유관속%형태
Rice%Shoot apical meristem%Young panicle%Vascular bundle%Morphology
通过体视显微镜及石蜡切片观察,研究了水稻幼穗分化过程中生长锥的形态以及维管束的生长变化。结果表明:水稻生长锥在幼穗分化第一期出现苞原基的分化,在第二期出现枝梗原基的分化,在第三期可见明显白色苞毛和颖花的发育,在第四期枝梗和颖花原基显著生长;在小穗发育过程中先后在枝梗末端分化出护颖原基、内外颖原基和雌雄蕊原基,在第四期花药中可见次级造孢细胞。发现在幼穗分化过程中幼穗主轴的维管束数目往上递减,穗茎节部位的大维管束能通过分离和合并来调节生长;在穗茎节出现的一种纤维状组织能够限制维管束的生长。
通過體視顯微鏡及石蠟切片觀察,研究瞭水稻幼穗分化過程中生長錐的形態以及維管束的生長變化。結果錶明:水稻生長錐在幼穗分化第一期齣現苞原基的分化,在第二期齣現枝梗原基的分化,在第三期可見明顯白色苞毛和穎花的髮育,在第四期枝梗和穎花原基顯著生長;在小穗髮育過程中先後在枝梗末耑分化齣護穎原基、內外穎原基和雌雄蕊原基,在第四期花藥中可見次級造孢細胞。髮現在幼穗分化過程中幼穗主軸的維管束數目往上遞減,穗莖節部位的大維管束能通過分離和閤併來調節生長;在穗莖節齣現的一種纖維狀組織能夠限製維管束的生長。
통과체시현미경급석사절편관찰,연구료수도유수분화과정중생장추적형태이급유관속적생장변화。결과표명:수도생장추재유수분화제일기출현포원기적분화,재제이기출현지경원기적분화,재제삼기가견명현백색포모화영화적발육,재제사기지경화영화원기현저생장;재소수발육과정중선후재지경말단분화출호영원기、내외영원기화자웅예원기,재제사기화약중가견차급조포세포。발현재유수분화과정중유수주축적유관속수목왕상체감,수경절부위적대유관속능통과분리화합병래조절생장;재수경절출현적일충섬유상조직능구한제유관속적생장。
Through observing the paraffin sections under stereoscopic microscope , we studied the morphology of shoot apical meristem ( SAM) in the course of young panicle differentiation as well as the growth changes of vascular bundles in rice .The results indicated that:in rice SAM, the differentiation of bract primordium appeared at the first stage of young panicle differentiation , the differentiation of branch primordium appeared at the second stage , the development of white bract hairs and glumous flowers was ob-vious at the third stage , and branch primordium and floret primordium grew observably at the fourth stage .In the process of spikelet development , glume primordium , inner glume primordium , outer glume primordium , pistils primordium and stamens primordium successively differentiated out from the end of branches , and the secondary sporogenous cells could be seen in the anthers in phaseⅣ.We found that:during the differentiation of young panicles , the number of vascular bundles became less in the higher position of the main branch of young panicles , the big vascular bundles in the neck-node of panicles could adjust their growth by segregation and recombination;a fibrous tissue in the neck-node of panicles could restrict the growth of vascular bundles .